Joe, can you clarify the Salmon thing? I'm not sure whether you're saying there are two or three phenotypes. If there are only two phenotypes, then salmon is not co-dominant, but completely dominant. If there are three phenotypes, it is codominant.
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The term
het strictly means that the paired genes are not the same. It has absolutely no dependence on outward appearances. If Salmon is codominant, then you have three genotypes:
- Normal (homozygous normal)
Salmon (heterozygous normal/salmon)
Super (homozygous salmon)
And the "Super" does NOT look like a Salmon, nor does it look like a Normal.
Codominance is more easily illustrated in roses:
If Red is dominant to White, here are the possibilities:
- Red/Red = RED
Red/White = RED (heterozygous)
White/White = WHITE
If Red and White are codominant to each other:
- Red/Red = RED
Red/White = PINK (heterozygous)
White/White = WHITE
Notice that you do NOT get red roses from crossing a red to a white rose. By the same token, you will NOT get "full bloodreds" by crossing bloodred to a non-carrier.
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The "diffuse pattern" appears to act like a codominant in many cases, and as a recessive in other cases. This is not unprecedented. In horses, the normal counterpart to the tobiano paint gene acts like a recessive in most cases, but occasionally it can be co-dominant. The same is true of the "roan" gene in horses. Pattern traits can vary in their expressiveness, even to the point where a "dominant" gene is almost completely undetectable when present.
Anyway, the argument for the "diffuse" pattern being a single (recessive or codominant) trait is very strong. IMO at least as strong as any of the other accepted mutants.
Look at the ancestry of my "bloodred" Mary, and it paints an interesting picture. She came from a Pewter crossed to an Anery het Bloodred and Amel. Now, let's look for the closest ancestor that could possibly be a "pure" (original line) bloodred:
The pewter had to have come from parents that were
both het for charcoal. The original bloods were not, so
neither of his parents were "originals." Best case scenario is that he came from double-hets, which would make his grandparents "Bloodred X Charcoal." This means that my bloodred, on her father's side, has at best a great-grandparent that was an original bloodred.
On the mother's side, we have pretty much the same deal. That is, an Anery het blood/amel couldn't have come directly from an "original" bloodred parent. Therefore, the "best" case scenario is that the Anery's grandparents were a Snow and a Bloodred, or something very similar. In this case, this gives my bloodred, at best, a great-grandparent on the mother's side, too.
(If Kathy got bored and actually wanted to look up her ancestry and see how close I guessed, she is a 1997 hatch from 92005 X 91113.)
How many selectively bred looks (Okeetee, Miami, Keys, Kisatchie) can be outcrossed and recovered, from unrelated lines, into F3s, from animals that don't necessarily show "the look" in the intermediate generations? Can you take some non-okeetee-looking normals that came from a single Okeetee ancestor and get any kind of consistent recovery of the Okeetee look? No way.
Also consider that my bloodred was bred to a pewter in 2000 and produced a clutch consisting entirely of offspring showing the same pattern. Two of those offspring were then bred to each other and produced "snow bloodred" offspring also expressing that "same" pattern.
This one family line has now carried the same pattern consistently and predictably through
at least five generations. Now, if you were to cross one of those "snow bloodred" individuals to Rich's bloodred lavender (which would mean they've been bred in very different directions for a good number of generations, at least six generations on one side) would anyone care to bet that the offspring will NOT show the "same pattern" as the proposed great, great, great, great grandparents did?
Would you make the same bet if I replaced "bloodred" with a "Miami phase" snow corn or an "Okeetee phase" lavender?